No, S3 doesn't cache any files locally. If a file were cached it's determined by your browser, who makes the decision based on a number of factors (such as HTTP headers, see [1]).
The reason the MP3 file plays in your browser is because your browser knows it can play the MP3 file and not just download it.
[1] https://developer.mozilla.org/en-US/docs/Web/HTTP/Headers/Cache-Control
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